Young and untested, Optimus Primal was a mere captain of an exploration ship when he and his crew were diverted off-course to pursue a stolen Predacon ship under the leadership of some crook named Megatron. The rest, they say, is (literally) history.
Though he does have a penchant for stuffy speeches, Optimus Primal is much more down-to-cybertron than his legendary namesake. He is committed to the Maximal codes of honor and moral judgment, but he does view them with some irreverence, and is not above finding loopholes or administering unorthodox solutions if necessary. ("Sometimes, crazy works", he says.) He is eternally loyal to his friends and respectful to his enemies. He is brave and selfless in battle. However, he is so committed to helping others that he can withdraw into a deep depression when he feels he has let them down. This is the only time he is self-absorbed.
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